Male flowerheads of Leucadendron spissifolium ssp. natalense in summer in the Mkhambathi Reserve on the Wild Coast, Eastern Cape, South Africa. [photo Colin Paterson-Jones ©]

A female plant of Leucadendron spissifolium ssp. natalense flowering in late spring in the Umtamvuna Reserve on the KwaZulu-Natal south coast, South Africa. [photo Colin Paterson-Jones ©].

A Male plant of Leucadendron spissifolium ssp. natalense flowering in summer in the Mkhambathi Reserve on the Wild Coast, Eastern Cape, South Africa. [photo Colin Paterson-Jones ©] 

A male plant of Leucadendron spissifolium ssp. spissifolium flowering in spring in the Kleinrivier Mountains, Western Cape, South Africa. [photo Colin Paterson-Jones ©]

A female plant of Leucadendron spissifolium ssp. spissifolium flowering in spring in the Elandsberg Mountains, Western Cape, South Africa. [photo Colin Paterson-Jones ©]

Leucadendron spissifolium ssp. fragrans male (left) and female (right) flowering in early summer on Robinson Pass in the Attaquaskloof Mountains, Western Cape, South Africa. [photos Colin Paterson-Jones ©]

Leucadendron spissifolium ssp. oribinum male (left) and female (right) flowering in early summer at Oribi Gorge in KwaZulu-Natal, South Africa. [photos Colin Paterson-Jones ©]